1 Bournemouth University, Faculty of Science and Technology, Department of Life and Environmental Sciences, Poole, Dorset BH12 5BB, UK; Harbor Branch Oceanographic Institute, Florida Atlantic University, Fort Pierce, FL 34946, USA.
2 Bournemouth University, Faculty of Science and Technology, Department of Life and Environmental Sciences, Poole, Dorset BH12 5BB, UK. Electronic address: gesteban@bournemouth.ac.uk.