1 Department of Ecology and Evolutionary Biology, Campus Box 334, University of Colorado Boulder, Boulder, Colorado 80304-0334 USA.
2 Department of Biological Sciences, 915 W. State Street, Purdue University, West Lafayette, Indiana 47907-0252 USA.
3 Department of Ecology and Evolutionary Biology, Campus Box 334, University of Colorado Boulder, Boulder, Colorado 80304-0334 USA Nancy.Emery@Colorado.edu.