1 School of Life Sciences and Biotechnology, Shanghai Jiaotong University, Shanghai 200240, China.
2 Key Laboratory of Urban Agriculture (South) Ministry of Agriculture and School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai 200240, China.
3 State Key Laboratory of Genetic Engineering and Collaborative Innovation Center for Genetics and Development, Institute of Plant Biology, School of Life Sciences, Fudan University, Shanghai 200438, China. Electronic address: hongquanyang@fudan.edu.cn.