1 Donghu Experimental Station of Lake Ecosystems, State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, 430072, Hubei, China.
2 Research Center of Lake Environment, Chinese Research Academy of Environmental Sciences, Beijing, 100012, China.
3 College of Fisheries, Huazhong Agricultural University, Wuhan, 430070, Hubei, China.
4 University of Chinese Academy of Sciences, Beijing, 100049, China.
5 Donghu Experimental Station of Lake Ecosystems, State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, 430072, Hubei, China. chenjun@ihb.ac.cn.