After publication of this article it was noticed there was an error in the Methods section under the subsection: Protein extraction and western blot analysis. The text including the error is as follows: "Denatured protein samples (15 μg) from each time-point were electrophoresed into 10 % SDS-PAGE gels (BioRad), transferred to PVDF membranes (BioRad) and incubated in primary antibodies overnight (Table 1)". Instead it should read: "…antibodies, C9ORF72 (1:500, Santa Cruz, sc-138763) and GAPDH (1:7000, Millipore), overnight." This error has since been updated in the article.