Two new species of Zospeum Bourguignat, 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae)

Adrienne Jochum; Anton J de Winter; Alexander M Weigand; Benjamín Gómez; Carlos Prieto

doi:10.3897/zookeys.483.9167

Two new species of Zospeum Bourguignat, 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae)

Zookeys. 2015 Feb 23:(483):81-96. doi: 10.3897/zookeys.483.9167. eCollection 2015.

Authors

Adrienne Jochum¹, Anton J de Winter², Alexander M Weigand³, Benjamín Gómez⁴, Carlos Prieto⁵

Affiliations

¹ Naturhistorisches Museum der Burgergemeinde Bern, CH-3005 Bern, Switzerland; Institute of Ecology and Evolution, University of Bern, CH-3012 Bern, Switzerland.
² Naturalis Biodiversity Center, P.O. Box 9517, 2300 RA Leiden, The Netherlands.
³ Department of Animal Ecology, Evolution and Biodiversity, Ruhr University Bochum, 44801 Bochum, Germany.
⁴ Department of Zoology, University of the Basque Country (UPV/EHU), 01006-Vitoria, Spain.
⁵ Department of Zoology and Animal Cell Biology, Faculty of Science and Technology, University of the Basque Country (UPV/EHU), 48080-Bilbao, Spain.

Abstract

Two new species of the genus Zospeum Bourguignat, 1856 are described from caves in the Sierra de Aitzgorri (Gipuzkoa) and the Sierra Salvada (Burgos) in Northern Spain. The taxa Zospeumvasconicum sp. n. and Zospeumzaldivarae sp. n. have recently, without a formal name, been included in a molecular study of worldwide members of the Carychiidae. In the present paper, the shell morphology and variation of these species is described and illustrated.

Keywords: Cave-dwelling species; conservation; ecology; microgastropoda; pseudo-cryptic species; shell variability; subterranean snail.