1 Department of Cell and Developmental Biology, The Institute for Regenerative Medicine and the Cardiovascular Institute, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA 19104, USA.
2 Department of Pediatrics and Biochemistry and Molecular Biology, Indiana University School of Medicine, Indianapolis, IN, USA.
3 Department of Medicine, Washington University School of Medicine, St Louis, MO 63110, USA.
4 Department of Neurology, Washington University School of Medicine, St Louis, MO 63110, USA. Electronic address: gutmannd@neuro.wustl.edu.