1 Centre for Biological Diversity and Scottish Oceans Institute, School of Biology, University of St. Andrews, St. Andrews, Fife, KY16 9TH, UK. maadd@st-andrews.ac.uk.
2 Department of Biology, University of Vermont, Burlington, VT 05405, USA.
3 School of Biology and Ecology, Sustainability Solutions Initiative, University of Maine, Orono, ME 04469, USA.
4 Centre for Biological Diversity and Scottish Oceans Institute, School of Biology, University of St. Andrews, St. Andrews, Fife, KY16 9TH, UK.